Nemo  2.4.0b
Simulate forward-in-time genetic evolution in a spatially explicit, individual-based stochastic simulator
LCEselection.h
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1 
30 #ifndef LCE_SELECTION_H
31 #define LCE_SELECTION_H
32 
33 #include <cmath>
34 #include "lifecycleevent.h"
35 #include "filehandler.h"
36 #ifdef HAS_GSL
37 #include <gsl/gsl_vector.h>
38 #include <gsl/gsl_matrix.h>
39 #endif
40 
41 class LCE_SelectionSH;
42 class LCE_SelectionFH;
43 //CLASS LCE_Selection_base
44 //
50 class LCE_Selection_base : public virtual LifeCycleEvent
51 {
54 
55  vector< TMatrix* > _selection_matrix;
56  vector< gsl_matrix* > _gsl_selection_matrix;
57  gsl_vector *_diffs,
65 
67  //double *_phe; // do we need this here?
68 
70 
75  vector<double> _mean_trait_value;
77 
80 
81  double _letheq;
82 
84  double _Fpedigree[5];
85 
88  double _FitnessFixModel[5];
89 
91 
92 
94  friend class LCE_SelectionSH;
95  friend class LCE_SelectionFH;
96  friend class LCE_Breed_Selection;
97 
98 protected:
99 
102  bool _is_local,
104 
106  double _fitness[5], _survival[5], _ind_cntr[5];
107 
110 
113 
115  vector< double > _selection_variance;
116 
118  double _eVariance;
120 
122  vector< string > _Traits;
123 
125  vector< unsigned int > _TraitIndices;
126 
128  vector< string > _SelectionModels;
129 
132 
133  vector< double (LCE_Selection_base::* ) (Individual*, unsigned int, unsigned int) > _getRawFitness;
134 
137  double (LCE_Selection_base::* _getFitness) (Individual*, unsigned int);
138 
142  void (LCE_Selection_base::* _setScalingFactor) (age_idx, unsigned int);
143 
147 
149 
152 
153 public:
154 
155  LCE_Selection_base ( );
156 
157  virtual ~LCE_Selection_base ( );
158 
159  bool setSelectionMatrix();
160  bool setLocalOptima ();
161  const TMatrix& getLocalOptima () const {return _local_optima;}
162  bool set_sel_model ();
163  bool set_fit_model ();
164  bool set_local_optima ();
166 
167  void set_std_rate_of_change();
168  void addPhenotypicSD (unsigned int deme, double *stDev);
169  void changeLocalOptima ();
170 
172  void resetCounters ();
173 
177  void setMeans (unsigned int tot_ind);
178 
184  void updateFitnessCounters (double fitness, unsigned int ped_class, bool survived);
185 
188  double getMeanFitness (age_idx age);
189 
193  double getMeanPatchFitness (age_idx age, unsigned int p);
194 
197  double getMaxFitness (age_idx age);
198 
202  double getMaxPatchFitness (age_idx age, unsigned int p);
203 
204 
207  double setMeanFitness (age_idx age) { return (_mean_fitness = getMeanFitness(age));}
208 
213  double getFitness (Individual* ind, unsigned int patch)
214  {
215  return (this->*_getFitness)(ind, patch);
216  }
217 
219  double getFitnessFixedEffect (Individual* offsprg, unsigned int patch, unsigned int trait)
220  {
221  return _FitnessFixModel[ offsprg->getPedigreeClass() ];
222  }
223 
225  double getFitnessDirect (Individual* offsprg, unsigned int patch, unsigned int trait)
226  {
227  return *(double*)offsprg->getTraitValue(trait);
228  }
230  double getFitnessUnivariateQuadratic ( Individual* ind, unsigned int patch, unsigned int trait);
231 
233  double getFitnessUnivariateLinear (Individual* offsprg, unsigned int patch, unsigned int trait);
234 
236  double getFitnessMultivariateGaussian (Individual* offsprg, unsigned int patch, unsigned int trait);
237 
239  double getFitnessUnivariateGaussian (Individual* offsprg, unsigned int patch, unsigned int trait);
240 
242  double getFitnessMultivariateGaussian_VE (Individual* offsprg, unsigned int patch, unsigned int trait);
243 
245  double getFitnessUnivariateGaussian_VE (Individual* offsprg, unsigned int patch, unsigned int trait);
246 
247 
252  double getFitnessAbsolute (Individual* ind, unsigned int patch);
253 
261  double getFitnessRelative (Individual* ind, unsigned int patch)
262  {
263  return getFitnessAbsolute(ind, patch) * _scaling_factor;
264  }
265 
269  void setScalingFactorLocal (age_idx age, unsigned int p);
270 
276  void setScalingFactorGlobal (age_idx age, unsigned int p);
277 
281  void setScalingFactorMaxLocal (age_idx age, unsigned int p);
282 
288  void setScalingFactorMaxGlobal (age_idx age, unsigned int p);
289 
294  void setScalingFactorForLinearSelection (age_idx age, unsigned int p);
295 
299  void setScalingFactorAbsolute (age_idx age, unsigned int p) { _scaling_factor = 1; }
300 
308  void doViabilitySelection (sex_t SEX, age_idx AGE, Patch* patch, unsigned int p);
309 
311  void checkChangeLocalOptima();
312 
315  virtual bool setParameters ();
316  virtual void execute ();
317  virtual void loadStatServices (StatServices* loader);
318  virtual void loadFileServices (FileServices* loader);
319  virtual bool resetParameterFromSource (std::string param, SimComponent* cmpt) {return false;}
320  virtual LifeCycleEvent* clone () {return new LCE_Selection_base();}
321  virtual age_t removeAgeClass () {return NONE;}
322  virtual age_t addAgeClass () {return NONE;}
323  virtual age_t requiredAgeClass () {return OFFSPRG;}
325 };
326 
327 
329 class LCE_SelectionSH : public EventStatHandler< LCE_Selection_base, LCE_SelectionSH >
330 {
331 
332  vector< vector< double > > _phenoTable;
334 
335 public:
336 
339  _table_set_gen(999999), _table_set_age(999999), _table_set_repl(999999)
340  {}
341 
342  virtual ~LCE_SelectionSH() {}
343 
344  virtual bool setStatRecorders (string& token);
345 
346  void addMeanPerPatch (age_t AGE);
347  void addVarPerPatch (age_t AGE);
348  void setDataTable (age_t AGE);
351 
352 
354  double getFitness (unsigned int i) {return _SHLinkedEvent->_fitness[i];}
355  double getSurvival (unsigned int i) {return _SHLinkedEvent->_survival[i];}
356  double getPedProp (unsigned int i) {return _SHLinkedEvent->_ind_cntr[i];}
357  double getMeanPatchFitness (unsigned int i, unsigned int int_agex);
358  double getMeanPatchFitness (unsigned int i);
359  double getVarPatchFitness (unsigned int i, unsigned int int_agex);
360 };
361 
362 class LCE_SelectionFH : public virtual EventFileHandler< LCE_Selection_base >
363 {
364 
365  public:
366 
368 
369  virtual ~LCE_SelectionFH() {}
370 
371  virtual void FHwrite();
372 
373  virtual void FHread(string& filename) {}
374 
375  void print(ofstream& FH, sex_t SEX, age_idx AGE, unsigned int i, Patch* patch, unsigned int ntraits);
376 };
377 
378 
379 #endif
380 
Template class for the LCEs StatHandler classes.
Definition: filehandler.h:254
Template class for the LCEs StatHandler classes.
Definition: stathandler.h:190
LCE_Selection_base * _SHLinkedEvent
Pointer to the linked LCE.
Definition: stathandler.h:193
A class to manage the files associated with each components of the simulation.
Definition: fileservices.h:52
This class contains traits along with other individual information (sex, pedigree,...
Definition: individual.h:49
void * getTraitValue(IDX T)
Accessor to the value (phenotype) of a particular trait.
Definition: individual.h:271
unsigned int getPedigreeClass()
Returns the pedigree class of the individual, as set during offspring creation.
Definition: individual.h:179
Composite LCE implementing breeding and viability selection on a given trait type.
Definition: LCEcomposite.h:238
Definition: LCEselection.h:363
virtual void FHwrite()
Definition: LCEselection.cc:1611
LCE_SelectionFH(LCE_Selection_base *event)
Definition: LCEselection.h:367
virtual void FHread(string &filename)
Definition: LCEselection.h:373
void print(ofstream &FH, sex_t SEX, age_idx AGE, unsigned int i, Patch *patch, unsigned int ntraits)
Definition: LCEselection.cc:1650
virtual ~LCE_SelectionFH()
Definition: LCEselection.h:369
StatHandler class for the LCE_Selection class.
Definition: LCEselection.h:330
double getVarPatchFitness(unsigned int i, unsigned int int_agex)
Definition: LCEselection.cc:1589
double getSurvival(unsigned int i)
Definition: LCEselection.h:355
unsigned int _table_set_age
Definition: LCEselection.h:333
double getMeanPatchFitness(unsigned int i, unsigned int int_agex)
Definition: LCEselection.cc:1562
void addVarPerPatch(age_t AGE)
Definition: LCEselection.cc:1457
unsigned int _table_set_repl
Definition: LCEselection.h:333
void setOffsprgTable()
Definition: LCEselection.h:350
double getMeanFitness()
Definition: LCEselection.h:353
void setAdultTable()
Definition: LCEselection.h:349
vector< vector< double > > _phenoTable
Definition: LCEselection.h:332
virtual ~LCE_SelectionSH()
Definition: LCEselection.h:342
virtual bool setStatRecorders(string &token)
Definition: LCEselection.cc:1365
void addMeanPerPatch(age_t AGE)
Definition: LCEselection.cc:1423
double getPedProp(unsigned int i)
Definition: LCEselection.h:356
double getFitness(unsigned int i)
Definition: LCEselection.h:354
unsigned int _table_set_gen
Definition: LCEselection.h:333
LCE_SelectionSH(LCE_Selection_base *event)
Definition: LCEselection.h:337
void setDataTable(age_t AGE)
Definition: LCEselection.cc:1491
Base class performing (viability) selection on an arbitrary trait.
Definition: LCEselection.h:51
double getFitnessUnivariateGaussian_VE(Individual *offsprg, unsigned int patch, unsigned int trait)
Returns the fitness of an individual following the Gaussian selection model with several traits under...
Definition: LCEselection.cc:896
int _selectTraitDimension
Number of quantitative traits under selection.
Definition: LCEselection.h:112
double getFitnessRelative(Individual *ind, unsigned int patch)
Returns the relative fitness of the individual, adjusted by a scaling factor.
Definition: LCEselection.h:261
bool setSelectionMatrix()
Definition: LCEselection.cc:545
double _fitness[5]
Fitness counters, one for each pedigree class.
Definition: LCEselection.h:106
double _base_fitness
Definition: LCEselection.h:100
double(LCE_Selection_base::* _getFitness)(Individual *, unsigned int)
Pointer to the function returning the individual fitness.
Definition: LCEselection.h:137
const TMatrix & getLocalOptima() const
Definition: LCEselection.h:161
virtual void loadFileServices(FileServices *loader)
Definition: LCEselection.cc:139
double getFitnessFixedEffect(Individual *offsprg, unsigned int patch, unsigned int trait)
Returns the fitness of an individual in the fixed selection model.
Definition: LCEselection.h:219
double getFitnessDirect(Individual *offsprg, unsigned int patch, unsigned int trait)
Returns the fitness of an individual following the direct selection model.
Definition: LCEselection.h:225
void setScalingFactorAbsolute(age_idx age, unsigned int p)
Resets the fitness scaling factor equal to one.
Definition: LCEselection.h:299
double _max_fitness
Definition: LCEselection.h:101
double _FitnessFixModel[5]
Absolute fitness values of the five pedigree class for the fixed selection model (lethal equivalents ...
Definition: LCEselection.h:88
double getMeanFitness(age_idx age)
Computes the mean fitness of the whole population for a given age class.
Definition: LCEselection.cc:945
vector< gsl_matrix * > _gsl_selection_matrix
Definition: LCEselection.h:56
double getFitnessUnivariateQuadratic(Individual *ind, unsigned int patch, unsigned int trait)
Quadratic fitness surface, approximates the Gaussian model for weak selection and/or small deviation ...
Definition: LCEselection.cc:839
void addPhenotypicSD(unsigned int deme, double *stDev)
Definition: LCEselection.cc:1312
void setScalingFactorMaxGlobal(age_idx age, unsigned int p)
Sets the fitness scaling factor equal to the inverse of the maximum population fitness value.
Definition: LCEselection.cc:1041
void doViabilitySelection(sex_t SEX, age_idx AGE, Patch *patch, unsigned int p)
Selectively removes individuals in the population depending on their fitness.
Definition: LCEselection.cc:1158
vector< double > _selection_variance
Patch-specific selection variance.
Definition: LCEselection.h:115
LCE_SelectionFH * _writer
Definition: LCEselection.h:151
vector< unsigned int > _TraitIndices
The indices of the traits under selection.
Definition: LCEselection.h:125
gsl_vector * _diffs
Definition: LCEselection.h:57
bool set_param_rate_of_change()
Definition: LCEselection.cc:740
void(LCE_Selection_base::* _setScalingFactor)(age_idx, unsigned int)
Pointer to the function used to set the fitness scaling factor when fitness is relative.
Definition: LCEselection.h:142
bool _is_absolute
Definition: LCEselection.h:103
double getMaxFitness(age_idx age)
Computes the maximum fitness value of the whole population for a given age class.
Definition: LCEselection.cc:979
TMatrix _linear_selection_offset
Definition: LCEselection.h:74
double _eVariance
Evironmental variance.
Definition: LCEselection.h:118
void setScalingFactorMaxLocal(age_idx age, unsigned int p)
Sets the fitness scaling factor equal to the inverse of the maximum local patch fitness value.
Definition: LCEselection.cc:1033
TMatrix _rate_of_change_local_optima
Definition: LCEselection.h:60
vector< double > _mean_trait_value
Definition: LCEselection.h:75
double _scaling_factor
Definition: LCEselection.h:101
bool _do_change_local_opt
Definition: LCEselection.h:61
bool _is_local
Definition: LCEselection.h:102
double getMeanPatchFitness(age_idx age, unsigned int p)
Computes the mean fitness in a given patch for a given age class.
Definition: LCEselection.cc:963
void checkChangeLocalOptima()
Check is rate of change in local optima has been set and must be applied.
Definition: LCEselection.cc:1105
virtual age_t addAgeClass()
Definition: LCEselection.h:322
virtual void execute()
Definition: LCEselection.cc:1124
double getFitness(Individual *ind, unsigned int patch)
Calls the fitness function according to the fitness model.
Definition: LCEselection.h:213
void setMeans(unsigned int tot_ind)
Computes the average fitness of each pedigree class.
Definition: LCEselection.cc:1200
virtual ~LCE_Selection_base()
Definition: LCEselection.cc:113
void resetCounters()
Resets the fitness counters.
Definition: LCEselection.cc:1189
double getFitnessMultivariateGaussian_VE(Individual *offsprg, unsigned int patch, unsigned int trait)
Returns the fitness of an individual following the Gaussian selection model with one trait under sele...
Definition: LCEselection.cc:912
gsl_vector * _res1
Definition: LCEselection.h:58
bool set_sel_model()
Definition: LCEselection.cc:304
double getFitnessMultivariateGaussian(Individual *offsprg, unsigned int patch, unsigned int trait)
Returns the fitness of an individual following the Gaussian selection model with one trait under sele...
Definition: LCEselection.cc:874
virtual age_t removeAgeClass()
Definition: LCEselection.h:321
double _ind_cntr[5]
Definition: LCEselection.h:106
virtual bool setParameters()
Definition: LCEselection.cc:178
unsigned int _set_std_rate_at_generation
Definition: LCEselection.h:63
double _mean_fitness
Definition: LCEselection.h:101
bool set_fit_model()
Definition: LCEselection.cc:246
virtual bool resetParameterFromSource(std::string param, SimComponent *cmpt)
Definition: LCEselection.h:319
LCE_Selection_base()
Definition: LCEselection.cc:46
LCE_SelectionSH * _stater
Definition: LCEselection.h:150
void setScalingFactorForLinearSelection(age_idx age, unsigned int p)
Sets the fitness scaling factor as the mean trait value in the patch to compute the fitness value in ...
Definition: LCEselection.cc:1050
int _std_rate_reference_patch
Definition: LCEselection.h:64
TMatrix _linear_selection_strength
Definition: LCEselection.h:73
void changeLocalOptima()
Definition: LCEselection.cc:1095
void(LCE_Selection_base::* _setNewLocalOptima)(void)
Pointer to the function used to change the local phenotypic optima or its rate of change.
Definition: LCEselection.h:146
double _Fpedigree[5]
Array of pedigree values used in the fixed-selection model.
Definition: LCEselection.h:84
TMatrix _local_optima
Definition: LCEselection.h:59
vector< double(LCE_Selection_base::*)(Individual *, unsigned int, unsigned int) > _getRawFitness
A vector containing pointers to fitness function related to each trait under selection.
Definition: LCEselection.h:133
bool set_local_optima()
Definition: LCEselection.cc:712
double _letheq
Definition: LCEselection.h:81
void setScalingFactorGlobal(age_idx age, unsigned int p)
Sets the fitness scaling factor equal to the inverse of the mean population fitness.
Definition: LCEselection.cc:1023
double getMaxPatchFitness(age_idx age, unsigned int p)
Computes the maximum fitness value in a given patch for a given age class.
Definition: LCEselection.cc:995
vector< TMatrix * > _selection_matrix
Definition: LCEselection.h:55
vector< string > _SelectionModels
The selection models associated with each trait under selection.
Definition: LCEselection.h:128
bool _rate_of_change_is_std
Definition: LCEselection.h:62
virtual LifeCycleEvent * clone()
Definition: LCEselection.h:320
double getFitnessUnivariateLinear(Individual *offsprg, unsigned int patch, unsigned int trait)
Returns the fitness of an individual following a Linear selection model with a single trait under sel...
Definition: LCEselection.cc:825
virtual age_t requiredAgeClass()
Definition: LCEselection.h:323
void setScalingFactorLocal(age_idx age, unsigned int p)
Sets the fitness scaling factor equal to the inverse of the mean local patch fitness.
Definition: LCEselection.cc:1015
double _survival[5]
Definition: LCEselection.h:106
vector< string > _Traits
The list of trait types under selection.
Definition: LCEselection.h:122
double getFitnessAbsolute(Individual *ind, unsigned int patch)
Returns the raw fitness of the individual, without adjustment (absolute fitness).
Definition: LCEselection.cc:933
virtual void loadStatServices(StatServices *loader)
Definition: LCEselection.cc:131
void set_std_rate_of_change()
Definition: LCEselection.cc:1228
double setMeanFitness(age_idx age)
Sets the _mean_fitness variable to the value of the mean population fitness.
Definition: LCEselection.h:207
double getFitnessUnivariateGaussian(Individual *offsprg, unsigned int patch, unsigned int trait)
Returns the fitness of an individual following the Gaussian selection model with several traits under...
Definition: LCEselection.cc:859
void updateFitnessCounters(double fitness, unsigned int ped_class, bool survived)
Updates the fitness and survival mean counters.
Definition: LCEselection.cc:1216
Base class of the Life Cycle Events, declares the LCE interface.
Definition: lifecycleevent.h:73
Second class in the metapopulation design structure, between the Metapop and Individual classes.
Definition: metapop.h:432
Interface to all basic components of a simulation (traits, life cycle events, pop,...
Definition: simcomponent.h:45
The Service class used to manage the StatHandler objects.
Definition: statservices.h:50
A class to handle matrix in params, coerces matrix into a vector of same total size.
Definition: tmatrix.h:50
sex_t
Sex types, males are always 0 and females 1!!
Definition: types.h:36
#define NONE
No age flag.
Definition: types.h:48
unsigned int age_t
Age class flags.
Definition: types.h:46
#define ADULTS
Adults age class flag (breeders).
Definition: types.h:54
#define OFFSPRG
Offspring age class flag.
Definition: types.h:50
age_idx
Array index of the age classes in the patch sizes and containers arrays.
Definition: types.h:41

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